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Glycophorin A （GYPA） and B （GYPB; this protein） are major sialoglycoproteins of the human erythrocyte membrane which bear the antigenic determinants for the MN and Ss blood groups respectively. In addition to the M or N and S or s antigens, that commonly occur in all populations, about 40 related variant phenotypes have been identified. These variants include the Miltenberger （Mi） complex and several isoforms of Stones （Sta）; also Dantu, Sat, Henshaw （He or MNS6）, Mg and deletion variants Ena, S-s-U- and Mk. Most of these are the result of gene recombinations between GYPA and GYPB.